TAXONOMY
This page is still being added to, but for the time being, I’ve put up a checklist of the 21 species of Agathis that are presently recognized in Farjon’s World Checklist & Bibliography of Conifers (2001) and in de Laubenfels’s account in Flora Malesiana, with their correct names and authors, and their types.
Note that some of the information in both these accounts is problematic. Note also that the Charles Moore Agathis collections in the Lindley Herbarium at Cambridge (CGE) appear to have been lost or destroyed.
Typification information comes primarily from Whitmore (Pl. Syst. Evol. 135 (1980): 41-69), Mabberley (Telopea 9: (2002): 743-754) and my own researches. Any corrections or queries would be extremely welcome.
Place of publication refers to the place of publication of the combination used here.
Place of typification refers to the publication in which the type of the species is designated, which may or may not be either the place of publication of this combination or the place of publication of the basionym. In some instances, where two syntypes are designated and one of these is later selected as lectotype, two references are given.
Synonymy will at some point be added.
**********
Agathis atropurpurea B.Hyland
TYPE: Hyland 5776
PLACE OF PUBLICATION: Brunonia 1: 109. (1978).
PLACE OF TYPIFICATION: Brunonia 1: 109. (1978).
BASIONYM: Agathis atropurpurea B.Hyland
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Queensland.
KNOWN TYPE LOCATIONS: K!, BRI
Agathis australis Lindl. in Loudon
TYPE: Illustration, in Lambert, Description of the Genus Pinus ed. 1, 2 (1824) 14, t.6. (see Whitmore 1980, Mabberley 2002).
PLACE OF PUBLICATION: Encycl. Pl.: 802. (1829).
PLACE OF TYPIFICATION: Mabberley (2002), Telopea 9: 752.
[The name Dammara australis D.Don in Lambert is typified by Whitmore (1980), in Pl. Syst. Evol. 135: 54]
BASIONYM: Dammara australis D.Don in Lambert, Description of the Genus Pinus ed. 1, 2 (1824) 14, t.6.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Northern North Island, New Zealand.
KNOWN TYPE LOCATIONS: Lambert’s Description of the Genus Pinus is quite widely distributed in botanical libraries, but no two copies are exactly the same. For a discussion, see: Renkema, H. W. & Ardagh, J. (1930). Aylmer Bourke Lambert and his ‘Description of the Genus Pinus’. Journal of the Linnean Society of London - Botany 48: 439-466.
Agathis borneensis Warb. (accidentally sunk in Farjon’s World Checklist.)
TYPE: Beccari 491
PLACE OF PUBLICATION: Monsunia 1: 184. (1900).
PLACE OF TYPIFICATION: Whitmore (1980), in Pl. Syst. Evol. 135: 55
BASIONYM: Agathis borneensis Warb.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Sumatra, Borneo, Peninsular Malaysia.
KNOWN TYPE LOCATIONS: K!, BM!, BO, P!, FI
Agathis corbassonii de Laub.
TYPE: de Laubenfels P419
PLACE OF PUBLICATION: Trav. Lab. For. Toulouse t. I vol. VIII art. V: 2. (1969).
PLACE OF TYPIFICATION: Trav. Lab. For. Toulouse t. I vol. VIII art. V: 2. (1969).
BASIONYM: Agathis corbassonii de Laub.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Northern New Caledonia.
KNOWN TYPE LOCATIONS: P!, K!
Agathis dammara (Lamb.) Rich. & A. Rich
The taxon to which this name refers is called A. celebica in both Flora Malesiana and Farjon’s World Checklist. The name Agathis dammara is mistakenly treated as the correct name for A. borneensis in Farjon’s World Checklist, probably as a result of a misinterpretation of an erroneous ‘Note’ on page 434 of the Agathis account in Flora Malesiana. For an explanation of this confusion, see the argument advanced by Veldkamp & de Laubenfels (1984) in Taxon 33: 337-347, and see also its rejection by the IAPT Committee for Spermatophyta, as being based on flawed premises, in Taxon 36: 734-762, and also my forthcoming paper.
TYPE: Illustration in Rumpf, Herbarium Amboinense 2 (1741), 174, t.57.
PLACE OF PUBLICATION: L. C. Richard & A. Richard, Comm. Bot. Conif. Cycad.: 83 (1826).
PLACE OF TYPIFICATION: Whitmore (1980), in Pl. Syst. Evol. 135: 57
BASIONYM: Pinus dammara Lambert, in Description of the Genus Pinus 1 (1803) 61, t.38.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Philippines, Sulawesi, Maluku.
KNOWN TYPE LOCATIONS: Herbarium Amboinense is widely distributed in botanical libraries.
Agathis endertii Meijer-Drees
TYPE: Netherlands Indies Forest Service bb21694
PLACE OF PUBLICATION: Bull. Jard. Bot. Buitenzorg, III, 16: 470 (1940).
PLACE OF TYPIFICATION: Bull. Jard. Bot. Buitenzorg, III, 16: 470 (1940).
BASIONYM: Agathis endertii Meijer-Drees
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Borneo
KNOWN TYPE LOCATIONS: BO
Agathis flavescens Ridl.
TYPE: Ridley 16023
PLACE OF PUBLICATION: Kew Bull. (1914): 332. (1914).
PLACE OF TYPIFICATION: Kew Bull. (1914): 332. (1914)
BASIONYM: Agathis flavescens Ridl.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. High mountaintops of Peninsular Malaysia.
KNOWN TYPE LOCATIONS: K!
Agathis kinabaluensis de Laub.
TYPE: de Laubenfels P625
PLACE OF PUBLICATION: Blumea 25: 535-537. (1979).
PLACE OF TYPIFICATION: Blumea 25: 535-537. (1979)
BASIONYM: Agathis kinabaluensis de Laub.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Mount Kinabalu, Crocker Range, Borneo
KNOWN TYPE LOCATIONS: A, K!, L, SAN
Agathis labillardierei Warb. (note correct orthography: see ICBN Art. 60.7 Ex. 11)
TYPE: La Billardi�re s.n. (in B-W, NOT destroyed during the 1939-1945 war, contra Whitmore 1980)
PLACE OF PUBLICATION: Monsunia 1: 183. (1900).
PLACE OF TYPIFICATION: Monsunia 1: 183. (1900)
BASIONYM: Agathis labillardierei Warb. (as ‘Agathis labillardieri’)
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Northern New Guinea, scattered westward from the Sepik Basin and northward from the Pegunungan Maoke, to the Vogelkopf and the islands of Waigeo, Japen and Biak.
KNOWN TYPE LOCATIONS: B-W
Agathis lanceolata Warb.
TYPE: Fournier & Sebert 60, p.p. (see Mabberley 2002)
PLACE OF PUBLICATION: Monsunia 1: 186. (1900).
PLACE OF TYPIFICATION: Mabberley, Telopea 9: 752.
BASIONYM: Agathis lanceolata Warburg
HABITAT AND GEOGRAPHIC RANGE: Rain forests and small patches of humid forest at tops of valleys. Southern and central New Caledonia.
KNOWN TYPE LOCATIONS: P!
Agathis lenticula de Laub.
TYPE: de Laubenfels P619
PLACE OF PUBLICATION: Blumea 25: 537-539. (1979).
PLACE OF TYPIFICATION: Blumea 25: 537-539. (1979).
BASIONYM: Agathis lenticula de Laub.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Borneo.
KNOWN TYPE LOCATIONS: A, K!, L, SAN
Agathis macrophylla (Lindl.) Mast.
TYPE: C. Moore s.n. (also interpreted as ‘No. 9’). (see Mabberley 2002)
PLACE OF PUBLICATION: J. Roy. Hort. Soc. 14: 197. (1892).
PLACE OF TYPIFICATION: Whitmore (1980), in Pl. Syst. Evol. 135: 61
Neither in the original description, nor in Mabberley (2002), nor in Whitmore (1980) is the species typified unambiguously onto to a numbered specimen, nor is this task performed by D�mmer (J. Roy. Hort. Soc. (1913) 39: 83), whose immensely useful study of the Lindley Herbarium at CGE sadly cannot be construed as an exercise in lectotypification. In any case, little of the Moore Agathis material at CGE seems to survive (it may have been destroyed during the 1950s). Whitmore (1980) lectotypifies the specimen on to C. Moore s.n.: given the ambiguities of the specimen numbering system adopted by Moore, this might well be the very same specimen as that listed by D�mmer and Mabberley as ‘No. 9’: certainly the specimen at K is unnumbered save for ‘9’ written in blue crayon on the type folder.
BASIONYM: Dammara macrophylla Lindley (1851), J. Hort. Soc. London 6: 271
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Vanikoro, Utupua, Nend� (Temotu Province, Solomon Islands); Erromango, Anatom, Tanna (Tafea, Vanuatu); Viti Levu, Vanua Levu (Fiji).
KNOWN TYPE LOCATIONS: †CGE Female cone material of holotype survives, mounted as if it were a diminutive stuffed owl, and has been seen; photograph at P!, K!
Agathis microstachya J.F.Bailey & C.T.White
TYPE: Mocatta s.n.
PLACE OF PUBLICATION: Contr. Queensland Fl. Bot. Bull. 18: 13. (1916).
PLACE OF TYPIFICATION: Contr. Queensland Fl. Bot. Bull. 18: 13. (1916).
BASIONYM: Agathis microstachya J.F.Bailey & C.T.White
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Queensland
KNOWN TYPE LOCATIONS: BRI, MEL
Agathis montana de Laub.
TYPE: Schmid 1420
PLACE OF PUBLICATION: Trav. Lab. For. Toulouse t. I vol. VIII art. V: 2. (1969)
PLACE OF TYPIFICATION: Trav. Lab. For. Toulouse t. I vol. VIII art. V: 2. (1969)
BASIONYM: Agathis montana de Laub.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Roches de Oua��me and summit ridge (> 1100 m) of the Massif du Pani�, northern New Caledonia.
KNOWN TYPE LOCATIONS: P!
Agathis moorei (Lindl.) Mast.
TYPE: C.Moore 8 (see Mabberley 2002)
PLACE OF PUBLICATION: J. Roy. Hort. Soc. 14: 197. (1892).
PLACE OF TYPIFICATION:
BASIONYM: Dammara moorei Lindley (1851), J. Hort. Soc. London 6: 271
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Northern and central New Caledonia.
KNOWN TYPE LOCATIONS: †CGE (nv), P! (photograph), ?K!, NSW (vide Mabberley 2002)
Agathis orbicula de Laub.
TYPE: de Laubenfels P614
PLACE OF PUBLICATION: Blumea 25: 540-541. (1979).
PLACE OF TYPIFICATION: Blumea 25: 540-541. (1979).
BASIONYM: Agathis orbicula de Laub.
HABITAT AND GEOGRAPHIC RANGE: Rain forests, Borneo.
KNOWN TYPE LOCATIONS: A, L, K!, SAN, SAR
Agathis ovata (Vieill.) Warb.
TYPE: Vieillard 1263 (see Mabberley 2002)
PLACE OF PUBLICATION: Monsunia 1: 186. (1900).
PLACE OF TYPIFICATION: de Laubenfels (1972), in Flore de la Nouvelle-Cal�donie et D�pendances 4: 133 (fide Mabberley 2002).
BASIONYM: Dammara ovata Vieillard, 1856, Ann. Sci. Nat. IV, 16: 56.
HABITAT AND GEOGRAPHIC RANGE: Largely confined to low maquis minier scrubland, though a very few scattered populations also occur in rain forest. Southern and central New Caledonia.
KNOWN TYPE LOCATIONS: P!
Agathis philippinensis Warb.
TYPE: Cuming 906
PLACE OF PUBLICATION: Monsunia 1: 185. (1900).
PLACE OF TYPIFICATION: Warburg (1900) in Monsunia 1: 185, Veldkamp & de Laubenfels (1984) in Taxon 33: 346.
BASIONYM: Agathis philippinensis Warburg
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Maluku, Sulawesi, Philippines.
KNOWN TYPE LOCATIONS: †B, BM!, L, CGE!, FI!, K!, P!, PNH, PR
Agathis robusta (C.Moore ex F.Muell.) F.M.Bailey
TYPE: T.W. Shepherd s.n. (MEL 258764)
PLACE OF PUBLICATION: Syn. Qld Fl. 498 (1883).
PLACE OF TYPIFICATION: Mabberley (2002), Telopea 9: 753
BASIONYM: Dammara robusta C.Moore ex F.Muell.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Queensland, Australia; Owen Stanley Range and central New Britain (Papua New Guinea).
KNOWN TYPE LOCATIONS: MEL
Agathis silbae de Laub.
TYPE: Askin 13156
PLACE OF PUBLICATION: Phytologia 61 (7): 448. (1987).
PLACE OF TYPIFICATION: Phytologia 61 (7): 448. (1987).
BASIONYM: Agathis silbae de Laub.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Mountaintops of Cumberland Range, western Espiritu Santo, Santo/Malo, Vanuatu.
KNOWN TYPE LOCATIONS: GH, K!, NY!, US!
Agathis spathulata de Laub.
TYPE: de Laubenfels P741
PLACE OF PUBLICATION: Flora Malesiana I, 10 (3): 435-436. (1988).
PLACE OF TYPIFICATION: Flora Malesiana I, 10 (3): 435-436. (1988)
BASIONYM: Agathis spathulata de Laub.
HABITAT AND GEOGRAPHIC RANGE: Rain forests. Eastern Highlands, Papua New Guinea.
KNOWN TYPE LOCATIONS: K!, L, US!